Eocene

Eocene (Gr. ἠώς, dawn, καινός, recent), in geology, the name suggested by Sir C. Lyell in 1833 for the lower subdivision of the rocks of the Tertiary Era. The term was intended to convey the idea that this was the period which saw the dawn of the recent or existing forms of life, because it was estimated that among the fossils of this period only 3½% of the species are still living. Since Lyell’s time much has been learned about the fauna and flora of the period, and many palaeontologists doubt if any of the Eocene species are still extant, unless it be some of the lowest forms of life. Nevertheless the name is a convenient one and is in general use. The Eocene as originally defined was not long left intact, for E. Beyrich in 1854 proposed the term “Oligocene” for the upper portion, and later, in 1874, K. Schimper suggested “Paleocene” as a separate appellation for the lower portion. The Oligocene division has been generally accepted as a distinct period, but “Paleocene” is not so widely used.

In north-western Europe the close of the Cretaceous period was marked by an extensive emergence of the land, accompanied, in many places, by considerable erosion of the Mesozoic rocks; a prolonged interval elapsed before a relative depression of the land set in and the first Eocene deposits were formed. The early Eocene formations of the London-Paris-Belgian basin were of fresh-water and brackish origin; towards the middle of the period they had become marine, while later they reverted to the original type. In southern and eastern Europe changes of sea-level were less pronounced in character; here the late Cretaceous seas were followed without much modification by those of the Eocene period, so rich in foraminiferal life. In many other regions, the great gap which separates the Tertiary from the Mesozoic rocks in the neighbourhood of London and Paris does not exist, and the boundary line is difficult to draw. Eocene strata succeed Cretaceous rocks without serious unconformity in the Libyan area, parts of Denmark, S.E. Alps, India, New Zealand and central N. America. The unconformity is marked in England, parts of Egypt, on the Atlantic coastal plain and in the eastern gulf region of N. America, as well as in the marine Eocene of western Oregon. The clastic Flysch formation of the Carpathians and northern Alps appears to be of Eocene age in the upper and Cretaceous in the lower part. The Eocene sea covered at various times a strip of the Atlantic coast from New Jersey southward and sent a great tongue or bay up the Mississippi valley; similar epicontinental seas spread over parts of the Pacific border, but the plains of the interior with the mountains on the west were meanwhile being filled with terrestrial and lacustrine deposits which attained an enormous development. This great extension of non-marine formations in the Eocene of different countries has introduced difficulties in the way of exact correlation; it is safer, therefore, in the present state of knowledge, to make no attempt to find in the Eocene strata of America and India, &c., the precise equivalent of subdivisions that have been determined with more or less exactitude in the London-Paris-Belgian area.

It is possible that in Eocene times there existed a greater continuity of the northern land masses than obtains to-day. Europe at that time was probably united with N. America through Iceland and Greenland; while on the other side, America may have joined Asia by the way of Alaska. On the other hand, the great central, mediterranean sea which stretched across the Eurasian continents sent an arm northward somewhere just east of the Ural mountains, and thus divided the northern land mass in that region. S. America, Australia and perhaps Africa may have been connected more or less directly with the Antarctic continent.

Associated, no doubt, with the crustal movements which closed the Cretaceous and inaugurated the Eocene period, there were local and intermittent manifestations of volcanic activity throughout the period. Diabases, gabbros, serpentines, soda-potash granites, &c., are found in the Eocene of the central and northern Apennines. Tuffs occur in the Veronese and Vicentin Alps—Ronca and Spelecco schists. Tuffs, basalts and other igneous rocks appear also in Montana, Wyoming, California, Oregon, Washington, Idaho, Colorado; also in Central America, the Antillean region and S. America.

It has been very generally assumed by geologists, mainly upon the evidence of plant remains, that the Eocene period opened with a temperate climate in northern latitudes; later, as indicated by the London Clay, Alum Bay and Bournemouth beds, &c., the temperature appears to have been at least subtropical. But it should be observed that the frequent admixture of temperate forms with what are now tropical species makes it difficult to speak with certainty as to the degree of warmth experienced. The occurrence of lignites in the Eocene of the Paris basin, Tirol and N. America is worthy of consideration in this connexion. On the other hand, the coarse boulder beds in the lower Flysch have been regarded as evidence of local glaciation; this would not be inconsistent with a period of widespread geniality of climate, as is indicated by the large size of the nummulites and the dispersion of the marine Mollusca, but the evidence for glaciation is not yet conclusive.

Organic Life of the Eocene Period.—As it has been observed above, the name Eocene was given to this period on the ground that in its fauna only a small percentage of living species were present; this estimation was founded upon the assemblage of invertebrate remains in which, from the commencement of this period until the present day, there has been comparatively little change. The real biological interest of the period centres around the higher vertebrate types. In the marine mollusca the most noteworthy change is the entire absence of ammonoids, the group which throughout the Mesozoic era had taken so prominent a place, but disappeared completely with the close of the Cretaceous. Nautiloids were more abundant than they are at present, but as a whole the Cephalopods took a more subordinate part than they had done in previous periods. On the other hand, Gasteropods and Pelecypods found in the numerous shallow seas a very suitable environment and flourished exceedingly, and their shells are often preserved in a state of great perfection and in enormous numbers. Of the Gasteropod genera Cerithium with its estuarine and lagoonal forms Potamides, Potamidopsis, &c., is very characteristic; Rostellaria, Voluta, Fusus, Pleurotoma, Conus, Typhis, may also be cited. Cardium, Venericardia, Crassatella, Corbulomya, Cytherea, Lucina, Anomia, Ostrea are a few of the many Pelecypod genera. Echinoderms were represented by abundant sea-urchins, Echinolampas, Linthia, Conoclypeus, &c. Corals flourished on the numerous reefs and approximated to modern forms (Trochosmilia, Dendrophyllia). But by far the most abundant marine organisms were the foraminifera which flourished in the warm seas in countless myriads. Foremost among these are the Nummulites, which by their extraordinary numerical development and great size, as well as by their wide distribution, demand special recognition. Many other genera of almost equal importance as rock builders, lived at the same time: Orthophragma, Operculina, Assilina, Orbitolites, Miliola, Alveolina. Crustacea were fairly abundant (Xanthopsis, Portunus), and most of the orders and many families of modern insects were represented.

When we turn to the higher forms of life, the reptiles and mammals, we find a remarkable contrast between the fauna of the Eocene and those periods which preceded and succeeded it. The great group of Saurian reptiles, whose members had held dominion on land and sea during most of the Mesozoic time, had completely disappeared by the beginning of the Eocene; in their place placental mammals made their appearance and rapidly became the dominant group. Among the early Eocene mammals no trace can be found of the numerous and clearly-marked orders with which we are familiar to-day; instead we find obscurely differentiated forms, which cannot be fitted without violence into any of the modern orders. The early placental mammals were generalized types (with certain non-placental characters) with potentialities for rapid divergence and development in the direction of the more specialized modern orders. Thus, the Creodonta foreshadowed the Carnivora, the Condylarthra presaged the herbivorous groups; but before the close of this period, so favourable were the conditions of life to a rapid evolution of types, that most of the great orders had been clearly defined, though none of the Eocene genera are still extant. Among the early carnivores were Arctocyon, Palaeonictis, Amblyctonus, Hyaenodon, Cynodon, Provivera, Patriofelis. The primitive dog-like forms did not appear until late in the period, in Europe; and true cats did not arrive until later, though they were represented by Eusmilus in the Upper Eocene of France. The primitive ungulates (Condylarths) were generalized forms with five effective toes, exemplified in Phenacodus. The gross Amblypoda, with five-toed stumpy feet (Coryphodon), were prominent in the early Eocene; particularly striking forms were the Dinoceratidae, Dinoceras, with three pairs of horns or protuberances on its massive skull and a pair of huge canine teeth projecting downwards; Tinoceras, Uintatherium, Loxophodon, &c.; these elephantine creatures, whose remains are so abundant in the Eocene deposits of western America, died out before the close of the period. The divergence of the hoofed mammals into the two prominent divisions, the odd-toed and even-toed, began in this period, but the former did not get beyond the three-toed stage. The least differentiated of the odd-toed group were the Lophiodonts: tapirs were foreshadowed by Systemodon and similar forms (Palaeotherium, Paloplotherium); the peccary-like Hyracotherium was a forerunner of the horse, Hyrochinus was a primitive rhinoceros. The evolution of the horse through such forms as Hyracotherium, Pachynolophus, Eohippus, &c., appears to have proceeded along parallel lines in Eurasia and America, but the true horse did not arrive until later. Ancestral deer were represented by Dichobune, Amphitragulus and others, while many small hog-like forms existed (Diplopus, Eohyus, Hyopotamus, Homacodon). The primitive stock of the camel group developed in N. America in late Eocene time and sent branches into S. America and Eurasia. The edentates were very generalized forms at this period (Ganodonta); the rodents (Tillodontia) attained a large size for members of this group, e.g. Tillotherium. The Insectivores had Eocene forerunners, and the Lemuroids—probable ancestors of the apes—were forms of great interest, Anaptomorphus, Microsyops, Heterohyus, Microchaerus, Coenopithecus; even the Cetaceans were well represented by Zeuglodon and others.

The non-placental mammals although abundant were taking a secondary place; Didelphys, the primitive opossum, is noteworthy on account of its wide geographical range.

Among the birds, the large flightless forms, Eupterornis, Gastornis, were prominent, and many others were present, such as the ancestral forms of our modern gulls, albatrosses, herons, buzzards, eagles, owls, quails, plovers. Reptiles were poorly represented, with the exception of crocodilians, tortoises, turtles and some large snakes.

The flora of the Eocene period, although full of interest, does not convey the impression of newness that is afforded by the fauna of the period. The reason for this difference is this: the newer flora had been introduced and had developed to a considerable extent in the Cretaceous period, and there is no sharp break between the flora of the earlier and that of the later period; in both we find a mixed assemblage—what we should now regard as tropical palms, growing side by side with mild-temperate trees. Early Eocene plants in N. Europe, oaks, willows, chestnuts (Castanea), laurels, indicate a more temperate climate than existed in Middle Eocene when in the Isle of Wight, Hampshire and the adjacent portions of the continent, palms, figs, cinnamon flourished along with the cactus, magnolia, sequoia, cypress and ferns. The late Eocene flora of Europe was very similar to its descendant in modern Australasia.